Malacologia

World’s Leading Malacological Journals Journal Impact Factor 1.592 (2012) Founded in 1961

EDITOR-IN-CHIEF

CHARLES LYDEARD e-mail

Editorial Office Malacologia CHARLES LYDEARD, Morehead State University, Department of Biology and Chemistry, 103 Lappin Hall, Morehead, KY  40351 U.S.A.

Business & Subscription Office Malacologia DAVID CAMPBELL, Box 7236, Department of Natural Sciences, Gardner-Webb University, Boiling Springs, NC  28017 U.S.A. e-mail

Copy Editor EUGENE COAN, Santa Barbara Museum of Natural History Santa Barbara, California, U.S.A. e-mail

Managing Editor CARYL HESTERMAN

“Honorary” Associate Editor JOHN B. BURCH, University of Michigan Ann Arbor, Michigan, U.S.A. e-mail

Composition Editor CLAUDIA WILKE, Giessen, Germany, e-mail

Advisory Editor THOMAS WILKE, Justus Liebig University, Giessen, Germany. e-mail

Malacologia is published by the INSTITUTE OF MALACOLOGY Board of Directors - U.S.A.:

President ELLEN STRONG, National Museum of Natural History, Washington D.C.

Vice President DIARMAID O FOIGHIL, University of Michigan, Ann Arbor

Secretary and Treasurer GEORGE M. DAVIS, West Falmouth, Massachusetts

Members

RÜDIGER BIELER, Field Museum, Chicago, Illinois

JOHN B. BURCH, University of Michigan, Ann Arbor

CAROLE S. HICKMAN, University of California, Berkeley

ALAN KOHN, University of Washington, Seattle

ELLEN STRONG, National Museum of Natural History, Washington D.C.

Copyright © 2014 by the Institute of Malacology ISSN: 0076-2997 (Print) ISSN: 2168-9075 (Online)