World’s Leading Malacological Journals

Journal Impact Factor 1.592 (2012)

Founded in 1961

EDITOR-IN-CHIEF Charles Lydeard

Editorial Office e-mail

Business & Subscription Office Malacologia P.O. Box 385 Haddonfield, New Jersey 08033, U.S.A. e-mail

Copy Editor EUGENE COAN Santa Barbara Museum of Natural History Santa Barbara, California, U.S.A. e-mail

Graphics Editor THOMAS WILKE Justus Liebig University Giessen, Germany e-mail

Associate Editor JOHN B. BURCH University of Michigan Ann Arbor, Michigan, U.S.A. e-mail

Managing Editor CARYL HESTERMAN Haddon Heights, New Jersey, U.S.A. e-mail

Composition Editor CLAUDIA WILKE Giessen, Germany e-mail

Malacologia is published by the INSTITUTE OF MALACOLOGY Board of Directors - U.S.A.:

President PAULA M. MIKKELSEN Paleontological Research Institution, Ithaca, New York

Vice President DIARMAID Ó FOIGHIL University of Michigan, Ann Arbor

Secretary and Treasurer GEORGE M. DAVIS West Falmouth, Massachusetts

Members RÜDIGER BIELER Field Museum, Chicago, Illinois

JOHN B. BURCH University of Michigan, Ann Arbor

CAROLE S. HICKMAN University of California, Berkeley

ALAN KOHN University of Washington, Seattle

ELLEN STRONG National Museum of Natural History, Washington D.C.

Copyright © 2014 by the Institute of Malacology ISSN: 0076-2997 (Print) ISSN: 2168-9075 (Online)