Staff Profile

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John Engel

Curator Emeritus

Negaunee Integrative Research Center

Additional Information


University of Wisconsin-Milwaukee, 1960-65.  B.S.

University of Wisconsin-Milwaukee, 1965-67.  M.S.

University of Michigan Biological Station, 1967.

Michigan State University, 1967-72.  Ph.D. 

Grants & awards

Senior Honors, 1964-65.

Delta Chi Sigma Honorary Science Fraternity (UW-M), 1964-67.

National Science Foundation Summer Fellowship, 1966.

University of Michigan award for study at University of Michigan Biological Station, 1967.

National Geographic Society Grant:  "Botanical Exploration and Collection of Bryophytes in Southern Chile," 1976.

National Science Foundation Grant:  "Systematics and Biogeography of Austro-Zelandic Hepaticae," 1976-79.

National Science Foundation Grant:  "The Systematics of Temperate and Subantarctic Australasian Lophocoleaceae,"  1981-85.

National Geographic Society Grant:  "Botanical Exploration and Collection of Bryophytes in Bolivia," 1982-83.

National Geographic Society Grant:  "Botanical Exploration and Collection of Bryophytes in Bolivia," 1985-86 (continuation of previous grant).

National Science Foundation:  "Curatorial Support for the Field Museum Herbarium," BSR-8801197 (7/15/88-12/31/91).

IllinoisMuseumServices:  Type Photo Restoration Grant (8/31/89-8/31/90).

National Science Foundation:  "Curatorial Support for the Field Museum Herbarium,"  BSR-9021960 (9/1/91-2/28/94).

National Geographic Society Grant:  "Investigation on the Hepatic flora of New Zealand," 1995.

National Geographic Society Grant:  "Interdisciplinary Investigations on the Hepatic and Agaric Biotas of New Zealand,"  1997.

National Geographic Society Grant:  "Studies on rare and threatened liverworts in the New Zealand botanical region."  2003.

National Geographic Society Grant:  "Biodiversity studies on the liverwort flora of the New Zealand Botanical Region."  2005.

National Geographic Society; "Liverworts in New Zealand:  Tree of Life and flora."  2006.

National Science Foundation Grant:  ATOL:  Collaborative Research -- Assembling the Liverwort Tree of Life:  A window into the evolution and diversication of early land plants.  2006-

GBIF Grant.  Towards a world checklist of liverworts.  (co-PI)

Work experience

Research Assistant, Michigan State University, 1967-71.

Postdoctoral Research Associate, 1971-72.

Donald Richards Assistant Curator of Bryology, Field Museum of Natural History, 1972-77.

Donald Richards Associate Curator of Bryology, Field Museum of Natural History, 1977-86.

Donald Richards Curator of Bryology, Field Museum of Natural History,  1986-2008

Served as Acting Chairman, Department of Botany, September 1987- May, 1988

Chairman, Department of Botany, Field Museum of Natural History, June, 1988 to Sept. 1993

Curatorial Affiliate in the Division of Botany, Peabody Museum of Natural History, Yale

Curator Emeritus, Department of Botany, Field Museum, 2009-



1.   A taxonomic monograph of the genus Balantiopsis (Hepaticae).  Nova Hedwigia 16: 83-130.  pl. 27 (1)-59 (33).  1968.

2.   Marsupidium in South America.  J. Hattori Bot. Lab. 34: 437-444.  pl. 1-2.  1971.  (with R. Grolle).

3.   The genus Andrewsianthus in South America and the Falkland Islands.  Bryologist 75: 325-334.  f. 1-64.  1972.

4.   Chiloscyphus hookeri n. sp. and nomenclatural changes in the genus Clasmatocolea.  J. Hattori Bot. Lab. 36: 150-156.  pl. 1-2.  1972 (1973).

5.   Metzgeria litoralis sp. nov. and Apometzgeria from southern South America.  Bryologist 76: 293-296.  f. 1-9.  1973.  (with Y. Kuwahara).

6.   On some tidal zone Hepaticae from South Chile, with comments on marine dispersal.  Bull. Torrey Bot. Club 100: 29-35.  1 tab.   1973.  (with R. M. Schuster).

7.   On the typification of Austrolophozia fuegiensis (Steph.) Schust. J. Hattori Bot. Lab. 37: 181-183.  1 tab.  1973.

8.   Austral Hepaticae I.  Pigafettoa Mass.  Bryologist 76: 511-515.  f. 1-9.  1974.  (with R. M. Schuster).

9.   Austral Hepaticae II.  Evansianthus, a new genus of Geocalycaceae.  Bryologist 76: 516-520.  f. 1-9.  1974.  (with R. M. Schuster).

10.   The Raymond E. Hatcher collection of Hepaticae and Anthocerotae from southern South America, with new taxa and notes on range extensions.  Bryologist 76: 528-535.  f. 1-18.  1974.

11.   Notes on the genus Hygrolembidium (Hepaticae).  Fieldiana, Bot. 36: 61-67.  f. 1-2.  1974.  

12.   A monograph of the genus Pseudocephalozia (Hepaticae).  J. Hattori Bot. Lab. 38: 665-701.  f. 1-17, tab. 1-2.  1974.  (with R. M. Schuster).

13.   Austral Hepaticae, V.  Studies on Schistochilaceae.  Phytologia 30: 241-250. 1975.  (with R. M. Schuster).

14.   Austral Hepaticae, IV.  Notes on Lophozia subg. Protolophozia Schust., with diagnosis of a new South American species.  J.  Bryol. 8: 465-474.  f. l.  1975.  (with R. M. Schuster).

15.   Austral Hepaticae III.  Stolonophora, a new genus of Geocalycaceae.  Fieldiana, Bot. 36: 111-124.  f. 1-6.  1975.  (with R.M. Schuster).

16.   Hepaticae and Anthocerotae collected by Dr. Harold E. Moore, Jr. in New Caledonia, Seychelles, Mauritius and Reunion in 1972.  Bryologist 78: 361-362.  1975.

17.   On the typification of Clasmatocolea obvoluta (Hook. f. & Tayl.) Grolle (Hepaticae).  Misc. Bryol. Lichenol. 7: 75.  1976.

18.   Metzgeria hamata Lindb., an illegitimate name of Hepaticae.  Lindbergia 3: 219-220.  f. 1.  1976.

19.   The southern South American Hepaticae and Anthocerotae collected by H. Roivainen in 1969-1970, with new taxa and notes on range  extensions.  Ann. Bot. Fenn. 13: 132-136.  1976.

20.   Austral Hepaticae VI.  Some new species and new combinations of taxa from southern Chile.  Bryologist 79: 514-515.  1976.

21.   Austral Hepaticae V.  The Schistochilaceae of South America.  J. Hattori Bot. Lab. 42: 273-423.  f. 1-45.  1977.   (with R. M. Schuster).

22.   Austral Hepaticae IX.  Anastrophyllum tristanianum, a new species from Tristan de Cunha.  Fieldiana, Bot. 38: 71-74.  f. 1-2.   1977.

23.   Index Hepaticarum Supplementum:  1974-1975.  Taxon 393-418.  1978.

24.   A taxonomic and phytogeographic study of Brunswick Peninsula (Strait of Magellan) Hepaticae and Anthocerotae.  Fieldiana, Bot. 41: i-viii, 1-319.  1978.

25.   Austral Hepaticae XI.  Lophocoleaceae:  New taxa, new combinations and re-alignments.  Phytologia 41: 309-312.  1979.

26.   Austral Hepaticae X.  A revision of Hepatostolonophora Engel et Schust., nom. nov. (Hepaticae).  J. Hattori Bot. Lab. 46: 91-108.  f. 1-8.  1979.

27.   Austral Hepaticae XII.  A new species of Clasmatocolea (Hepaticae) from Tasmania.  Bryologist 83: 220-223.  f. 1-19.  1980.

28.   A monograph of Clasmatocolea (Hepaticae).  Fieldiana, Bot. N. S. 3: i-viii, 1-229.  f. 1-61, pl. 1-29.  1980.

29.   Austral Hepaticae XIII.  Two new genera of Geocalycaceae (Lophocoleaceae).  Phytologia 47: 309-312.  1981.  (with R. M.  Schuster).

30.   Austral Hepaticae XV.  Brevianthaceae, fam. nov. and Brevianthus, gen. nov. from Tasmania.  Phytologia 47: 317-318.  1981.  (with R. M. Schuster).

31.   Index Hepaticarum Supplementum:  1976-1977.  Taxon 30: 518-537.  1981.

32.   Austral Hepaticae XVII.  Pachyschistochila Schust. et Engel, gen. nov. Phytologia 50: 177-180.  1982.  (with R. M. Schuster)

33.   Hepaticopsida.  In:  Parker, S. P. (ed.).  Synopsis and Classification of Living Organisms.  Vol. 1, pp. 271-304.  McGraw-Hill, New York.  1982.

34.   Anthocerotopsida.  In:  Parker, S. P. (ed.)  Synopsis and Classification of Living Organisms.  Vol. 1, pp. 304-305.  McGraw-Hill, New York.  1982.

35.   Haplomitrium monoicum, a remarkable new species of Calobryales (Hepaticae) from New Caledonia, together with a reclassification  of Subg. Haplomitrium.  Ann. Missouri Bot. Gard. 68: 668-676,  figs. 1-8.  1982 [1981].

36.   Austral Hepaticae XVI.  Gondwanalandic Leptoscyphoideae (Geocalycaceae)  Lindbergia 8: 65-74, figs. 1-3.  1982.  (with R. M. Schuster).

37.   Austral Hepaticae XV.  Brevianthaceae:  A monotypic family endemic to Tasmania.  Bryologist 85: 375-388, figs. 1-39.  1982 [1983].   (with R. M. Schuster).

38.   Austral Hepaticae XVIII.  Studies toward a revision of Telaranea subg. Neolepidozia (Lepidoziaceae).  Fieldiana Bot. N.S. 14: i- v, 1-7, figs. 1-3.  1983.  (with R. M. Schuster).

39.   Review of Karen S. Renzaglia, A comparative developmental investigation of the gametophyte generation in the Metzgeriales  (Hepatophyta).  Bryophytorum Bibliotheca 24: i-x, 1-253.  Cramer, 1982.  Bryologist 87:  93-95. 1984.

40.   Botanical exploration and collection of bryophytes in southern Chile.  National Geographical Research Reports Vol. 16: 239-244.  1984.

41.   An overview and evaluation of the genera of Geocalycaceae Subfamily Lophocoleoideae (Hepaticae).  Nova Hedwigia 39: 385-463, figs. 1-10.  1984.  (with R. M. Schuster)

42.   Index Hepaticarum Supplementum:  1978-1979.  Taxon 33: 761-779.  1984.

43.   Leptophyllopsis and LeptoscyphusIn: Geissler, P. & H. Bischler (ed.), Index Hepaticarum.  Vol. 10: 65-70.  1985.  J. Cramer.  Vaduz, Germany.

44.   Austral Hepaticae V (2).  Temperate and subantarctic Schistochilaceae of Australasia.  J. Hattori Bot. Lab. 58: 255-539, figs. 1-76.  1985.  (with R. M. Schuster)

45.   Psiloclada major (Schust.) Engel, comb. nov. (Hepaticae) from New Zealand.  Phytologia 58: 324.  1985. 

46.   An annotated checklist of Philippine Hepaticae.  J. Hattori Bot. Lab. 60: 283-355.  1986. (with Benito Tan).

47.   Austral Hepaticae XX.  New species of Hygrolembidium (Lepidoziaceae).  Phytologia 62: 9-12.  1987.

48.   Miscellanea Hepaticologica Paleotropica.  I.  Pp. 533-534.  In:  Buck, W. (ed.), Bryostephane Steereana:  A Collection of Bryological Papers Presented to William Campbell Steere on the Occasion of his 80th Birthday.  New York Bot. Garden.  1987.

49.   A Monograph of Lepidoziaceae subfam. Lembidioideae (Hepaticae).  Journal of the Hattori Botanical Laboratory 63: 247-355.  1987.  (with R.M. Schuster).

50.   Review of:  G.A.M. Scott, Southern Australian Liverworts.  Pp. i-ix, 1-216.  Australian Flora and Fauna Series no. 2, Bureau of Flora and Fauna, Australian Government Publishing Service, Canberra.  1985. Taxon 36: 774-775.  1987.

51.   Studies of New Zealand Hepaticae.  1-6.  Brittonia 40: 200-207, f. 1-18.  (with R.M. Schuster).  1988.

52.   Bryological Contributions.  Presented in Celebration of the Distinguished Scholarship of Rudolf M. Schuster.  Beih. Nova Hedwigia 90: 1-397. (J. Engel, ed; with S. Hattori as consulting ed.)  1988.

53.   The taxonomic position of Apotomanthus (Hepaticae).  Pp. 203-221, figs. 1-4.  In: Engel, J. and S. Hattori (eds.), Bryological Contributions Presented in Celebration of the Distinguished Scholarship of Rudolf M. Schuster.  Beih. Nova Hedwigia 90.  1988.

54.   Index Hepaticarum Supplementum:  1980-1981.  Taxon 38: 414-439.  1989.

55.   PachyglossaIn:  Geissler, P. & H. Bischler (ed.), Index Hepaticarum.  Vol. 11:  55-56.  J. Cramer, Berlin.  1989.

56.   Saccogyna, Saccogynidium, Schistochila, Schistochilaster, Tetracymbaliella and XenocephaloziaIn:  Geissler, P. & H. Bischler (ed.), Index Hepaticarum.  Vol. 12:  J. Cramer, Berlin.  1990.

57.   A preliminary study on the affinities of Philippine, Bornean and New Guinean hepatics.  Tropical Bryology 2: 265-272.  (with B. Tan).  1990.

58.   Index Hepaticarum Supplementum:  1982-1983.  Taxon 39:  245-254.  1990.

59.   Falkland Island (Islas Malvinas) Hepaticae and Anthocerotae:  A taxonomic and phytogeographic study.  Fieldiana, (Bot.) N.S. 25: i-viii, 1-209, figs. 1-88. 1990.

60.   Studies on Geocalycaceae.  I.  The taxonomic position of Chiloscyphus amplectens (Mitt.) Engel & Schust. together with refinements in Heteroscyphus Schiffn.  Journal Hattori Botanical Laboratory 68:  303-315.  1990.

61.   Studies on Geocalycaceae (Hepaticae).  II.  Stolonivector, a new genus from New Zealand.  J. Hattori Bot. Lab. 69: 79-86.  1991.

62.   Studies on Geocalycacaeae (Hepaticae).  III. On the typification of Jungermannia multipenna Hook. f. & Tayl.  Bryologist 94: 22-24.  1991.

63.   Studies on Tasmanian Hepaticae I.  The genus Adelanthus.  Pp. 7-11.  In:  M.R. Banks et al. (eds.). Aspects of Tasmanian Botany.  A tribute to Winifred Curtis.  Royal Society of Tasmania, Hobart.  1991. Pap. & Proc. Roy. Soc. Tasmania.

64.   Studies on Geocalycaceae (Hepaticae).  IV.  Lamellocolea, a new genus of Leptoscyphoideae from New Zealand.  J. Hattori Bot. Lab. 70: 63-78, f. 1-4.  1991.

65.   Studies on Geocalycaceae (Hepaticae).  V.  Comments on the Leucophyllae complex of Chiloscyphus.  Bryologist 94: 311-314.  1991.

66.   Studies on Geocalycaceae (Hepaticae).  VI.  Comments on Leptoscyphus in Australasia, together with refinements in Clasmatocolea.  Bryologist 94: 435-437.  191.

67.  Index hepaticarum supplementum:  1984-1985.  Taxon 41: 253-306.  1992.

68.   Studies on Geocalycaceae (Hepaticae).  VII.  Subspecific differentiation of Chiloscyphus semiteres together with further refinements in Chiloscyphus (s. lat.).  Contr. Univ. Mich. Herb. 18: 107-111, fig. 1.  1992.

69.   Studies on Geocalycaceae (Hepaticae).  VIII.  A revision of Chiloscyphus subg. Notholophocolea (Schust.) Engel & Schust.  J. Hattori Bot. Lab. 72: 105-115, f. 1-3.  1992.

70.   Review of Howard Crum, Liverworts and hornworts of Southern Michigan.  Univ. of Michigan Herb.  233 pp. 1991.  Michigan Botanist.  Michigan Botanist 30:  134-135.

71.   Review D. J. Galloway.  Studies in Pseudocyphellaria (Lichens) III.     The South American Secies.  Bibliotheca Lichenologica 46: 1-275.  Cramer, 1992.  Bryologist 96:  178-179. 1993.

72.   Studies on Geocalycaceae (Hepaticae).  IX.  Chiloscyphus perpusillus    (Hook. f. & Tayl.) Engel:  A study in novel asexual reproductive strategies.  Nova Hedwigia 56: 241-246, f. 1-2.  1993.

73.   Index Hepaticarum Supplementum:  1986-1987.  Taxon 42: 373-391.  1993.

74.  Studies on Geocalycaceae.  IX.  Chiloscyphus hattorii Engel, a new species from New Zealand, together with nomenclatural refinements in Australasian Heteroscyphus and Leptoscyphus.  J. Hattori Bot. Lab. 74:  29-33, f. 1-2.  1993.

75.   Studies of New Zealand Hepaticae.  7. The status of Steereomitrium.  Bryologist 97: 63-66, f. 1.  1994.  (with R. M. Schuster)

76.   Studies of New Zealand Hepaticae.  8-13.  Bazzania and Acromastigum.   Bryologist 97: 313-320, f. 1-3.  1994.  (with G. Merrill)

77.   Austral Hepaticae.  XXI.  Paracromastigum fiordlandiae, sp. nov. and the delimitation of Paracromastigum and Hyalolepidozia (Lepidoziaceae).  Brittonia 48: 165-173, f. 1.  1996.   (with R. M. Schuster)

78.   Austral Hepaticae 23.  New taxa and new combinations in Telaranea Spruce ex Schiffn. (Lepidoziaceae).   Phytologia 79: 250-253.  June, 1996 (1995).   (with G. Merrill)

79.   Studies of New Zealand Hepaticae.  14-19.  Kurzia and Lepicolea.  J. Hattori Bot. Lab. 80: 217-231, f. 1-3.  1996 (October).  (with G. Merrill)

80.   Austral Hepaticae. 22.  The genus Balantiopsis in New Zealand, with observations on extraterritorial taxa and a phylogeny of Balantiopsis and the Family Balantiopsaceae (Jungermanniales).   Fieldiana, Bot. 37: i-iii, 1-62, f. 1-18.  1997.   (with G. Merrill)

81.   Austral Hepaticae  XXIV.  A Revision of Isotachis Mitt. (Balantiopsaceae:  Isotachidoideae) in New Zealand.  J. Hattori Bot. Lab. 83: 187-227, f. 1-15. 1997.   (with R. M. Schuster)

82.   Review of Martin J. Wigginton and Riclef Grolle, Catalogue of the Hepaticae and Anthocerotae of Sub-Saharan Africa.  Bryophytorum Bibliotheca 50: 1-267. J. Cramer, D-70176 Stuttgart.  1996.  Bryologist 100: 556-557.  997.

83.   Studies on Geocalycaceae (Hepaticae). X. New taxa and new combinations in Chiloscyphus Corda for Australasia.  Phytologia 83: 42-46.  ?May, 1998 (1997).

84.   Austral Hepaticae XXV.  Krunodiplophyllum Grolle and a revision of the Australasian species of Diplophyllum (Dum.) Dum. (Scapaniaceae, Jungermanniales).  J. Hattori Bot. Lab. 84: 241-283. f. 1-9.  1998.  (with G. Merrill)

85.   Austral Hepaticae. 27.  The genus Anastrophyllum (Spruce) Steph. (Jungermanniales) in Australasia, with a synopsis of Austral taxa.  Journal of Bryology 20: 371-388. f. 1-3.  1998 (with J. Braggins)

86.   Studies on Geocalycaceae (Hepaticae). XI. Supraspecific new taxa and new combinations in Chiloscyphus Corda for Australasia.  Novon 9: 22-24.  1999.

87.   Austral Hepaticae. 26. The identity, taxonomic position and ecology of Trichocolea julacea Hatcher (Trichocoleaceae).  Novon 9: 25-28. fig. 1.  1999.

88.   Austral Hepaticae 28. Plagiochila bazzanioides Engel & Merrill, a remarkable new species of Plagiochilaceae from New Zealand.  Novon 9: 29-31. fig. 1.  1999. (with G. Merrill)

89.   Austral Hepaticae 29.  More new taxa and combinations in Telaranea (Lepidoziaceae) and a new name for Frullania caledonica (Schust.) Schust. (Frullaniaceae) from New Caledonia.   Novon 9: 339-344, f. 1-3.  1999. (with G. Merrill)

90.   Austral Hepaticae. 30.  A critical new species of Triandrophyllum (Herbertaceae) from New Zealand.  Haussknechtia Beihefta 9: 115-119, f. 1-2.  1999. 

91.   Austral Hepaticae 31.  Two new species of Plagiochila (Dum.) Dum. (Plagiochilaceae) from New Zealand. J. Hattori Bot. Lab. 87: 295-300, f. 1-2.  1999.  (with G. Merrill)  (Nov.)

92.   Austral Hepaticae 33.  Paracromastigum succulentum (Sim) Engel & Merr., comb. nov.  (Lepidoziaceae, Jungermanniales).  Bryologist 104: 151-153, f. 1.  2001.  (with G. L. Merrill)

93.   Studies on Geocalycaceae.  XII.  Heteroscyphus mononuculus Engel, a new species of Hepaticae from New Zealand.  J. Hattori Bot. Lab. 90: 241-244, f. l.  2001.

94.   Austral Hepaticae 32. Revision of the genus Lepidozia (Hepaticae) for New Zealand.  Fieldiana, Bot. 42: 1-107, f. 1-39.  2001.  (with R. M. Schuster)

95.   Austral Hepaticae 34.  The sporophyte of Neogrollea Hodgs. and the taxonomic position of Neogrolleaceae (Schust.) Engel & Braggins comb. & stat. nov.  J. Hattori Bot. Lab. 91: 173-204, f. 1-17.  2001.  (with J. Braggins)

96.   (1544)  Proposal to conserve Telaranea against Arachniopsis (Hepaticae).  Taxon 51: 571-572.  2002.  (with G. Merrill)

97.   Studies on Geocalycaceae (Hepaticae).  XIII.  The genus Stolonivector Engel, including a new species from New Zealand.  J. Hattori Bot. Lab. 93: 69-77, f. 1-3.  2003.

98.   Studies on Geocalycaceae XIV.  Physotheca J. J. Engel & Gradst., a new genus of Hepaticae from Ecuador, belonging to a new subfamily, Geocalycaceae subfam. Physothecoideae J. J. Engel & Gradst.  Taxon. 52: 763-773, f. 1-3.  2003.  (with S. Rob Gradstein)

99.   Collecting Liverworts Down Under.  In the Field 75: 18.  2004.  (with Matt von Konrat)

100.  Studies on Geocalycaceae.  XV. Chiloscyphus aperticaulis Engel, an interesting new species of Hepaticae from New Zealand.  J. Hattori Bot. Lab. 95: 229-234, f. 1-2.  2004 (Jan.)

101.  Austral Hepaticae 36.  A new species of Lepidozia from New Zealand, together with an assessment of Subg. Austrolepidozia (Schust.) Schust.  J. Hattori Bot. Lab. 96: 273-279, f. 1-3.  2004.

102.  Austral Hepaticae. 35.  A taxonomic and phylogenetic study of Telaranea (Lepidoziaceae), with a monograph of the genus in temperate Australasia and commentary on extra-Australasian taxa.  Fieldiana NS 44: i-iv, 1-265, f. 1-86, tab. 1-4.  (with G.L. Smith Merrill).  2004

103.Austral Hepaticae 37.  The sporophyte of Megalembidium Schust., together with a re-evaluation of the taxonomic position of the genus.  J. Hattori Bot. Lab. 97: 91-96, f. 1-8.  2005  (with J. Braggins).

104.Austral Hepaticae 39.  Kurzia moniliformis, an interesting new species of Hepaticae from New Zealand, belonging to a new section, Kurzia sect. Moniliformes Engel.  Cryptogamie, Bryologie 26: 73-78, f. 1.  2005.

105.Austral Hepaticae 38.  The gynoecium of Trichotemnoma (Steph.) Schust., together with a re-evaluation of the taxonomic position of Trichotemnomaceae Schust. and comments on family endemism in south temperate areas.  Nova Hedwigia 80: 367-385, f. 1-3.  2005.

106.Are Mylia and Trabacellula (Hepaticae) related?  Unsuspected links revealed by cell wall morphology, with the transfer of Mylia anomala to a new genus (Leiomylia J. J. Engel & Braggins) of Jungermanniaceae.  Taxon 54: 665-680, f. 1-7.  2005.

107.Rare and endangered liverworts:  A new area of conservation in New Zealand.  Australasian Plant Conservation 14: 17-19.  2005. (M.J. von Konrat, D.S. Glenny, P.J. de Lange, J.J. Engel, J.E. Braggins and M.A.M. Renner)

108.Austral Hepaticae 40.  Tritomaria (Jungermanniaceae subfam. Lophozioideae) new to the south temperate, together with a new subspecies, T. exsecta subsp. novaezelandiae subsp. nov.  Bryologist 109: 60-67, f. 1-2.  2006.

109.Austral Hepaticae 41.  Bazzania exempta Engel, an interesting new species from New Zealand, belonging to a new section, Bazzania sect. Exemptae Engel.  J. Hattori Bot. Lab. 99: 197-205, f. 1-3.  2006.

110.Austral Hepaticae 42.  The Austral species of Mnioloma (Calypogeiaceae), together with a new species, Mnioloma novaezelandiae n. sp.  Cryptogamie, Bryologie:  27: 111-117.  f. 1-2.  2006.

111.Index of Hepatics 1974-2000.  Pp. 1-368. Hattori Botanical Laboratory. Nichinan, Miyazaki, Japan.  2006.  (with M. Crosby)

112.Goodbye or welcome Gondwana? -- insights into the phylogenetic biogeography of the leafy liverwort Plagiochila with a description of Proskaurera, gen. nov.  (Plagiochilaceae, Jungermanniales.  Pl. Syst. Evol. 258: 227-250.  Heinrichs, J., M. Lindner, H. Groth, J. Hentschel, K. Feldberg, C. Renker, J.J. Engel, M. von Konrat, D.G. Long and H. Schneider.  2006.

113.Studies of New Zealand Hepaticae.  20-38.  A miscellanea of new taxa and combinations.  Novon 17: 310-314.  2007.  (10 Sept.)

114.Austral Hepaticae 43.  Castanoclobos, a new genus of Trichocoleaceae from New Zealand.  Novon 17: 424-428, f. 1-2.  2007.    J.J. Engel and D. Glenny  (3 Dec.)

15.Studies on Lophocoleaceae XVII.  On Leptophyllopsis R.M. Schust., with particular reference to L. laxus (Mitt.) R.M. Schust., together with a new subfamily, Lophocoleaceae subfam. Leptophyllopsoideae J.J. Engel  Nova Hedwigia 417-428, f. 1-5.  2007.

116.Studies on Lophocoleaceae XVI.  Chiloscyphus anisolobus, an interesting new species from New Zealand.  Bryologist 111: 118-123, f. 1-2.  2008.      J.J. Engel and D. Glenny

117.Studies of New Zealand Hepaticae.  39.  Dinckleria Trev. an older name for Proskauera Heinrichs and Engel.  Cryptogamie, Bryologie 29: 193-194.  2008.  J.J. Engel and J. Heinrichs

118.A flora of the liverworts and hornworts of New Zealand.  Vol. 1.  Monographs in Systematic Botany 110:  1-882.  Missouri Botanical Garden Press.  St. Louis, Missouri.  2008.  J.J. Engel and D. Glenny

119.Austral Hepaticae 44.  A revision of Marsupidium Mitt. for New Zealand.  Nova Hedwigia 87: 277-313, f. 1-10.  2008.  J.J. Engel and D. Glenny

120.The systematic identity of Chiloscyphus trichocoleoides, a new liverwort species from New Zealand, uncovered by morphological and molecular evidence.  J. Bryol. 31: 93-105, f. 1-8.  2009.  Glenny, D., J.J. Engel and X. He-Nygrén

121.Studies on Lophocoleaceae (Hepaticae).  XVIII.  Stolonivector Engel in New Zealand, including two new species, together with comments on generic endemism of Austral Hepaticae.  Nova Hedwigia 88: 335-346, f. 1-3.  2009

122.Early land plants today:  Global patterns of liverwort diversity, distribution, and floristic knowledge.  Pp. 425-438. In: Mohamed, H., B.B. Bakar, A. N. Boyce and P. K. Y. Lee (eds.).  Bryology in the New Millennium.  City Reprographic Services, Kuala Lumpur.  von Konrat, M., A. Hagborg, L. Söderstrom, J. Mutke, M. Renner, R.R. Gradstein, J. Engel, R.-L. Zhu and J. Pickering.  2008.

123.Austral Hepaticae 46.  The identity of Plagiochila retrospectans (Nees ex Spreng.) Lindenb. and P. fuscella (Hook. f. & Tayl.) Gottsche, Lindenb. & Nees, two misunderstood Australasian species.  Nova Hedwigia 89: 287-301, f. 1-5.  J.J. Engel and G. L. Merrill.  2009.  (ca. Nov.)

124.Early land plants today:  Index of liverworts & hornworts 2006-2008. Cryptogamie, Bryologie 31: 3-30.  von Konrat, M., L. Söderstrom, A. Hagborg, M. Crosby and J. Engel.  2010.

125.Austral Hepaticae 45. A monograph of the genus Chiloscyphus Corda (Lophocoleaceae) for Australasia.  Fieldiana, Bot. N.S. 48: i-vi, 1-209, f. 1-81.  (4 Mar.)  2010.

126.Studies on Lophocoleaceae. XIX. The systematic identity of Cyanolophocolea R.M. Schust., an intriguing liverwort from New Zealand and Australia, based on morphological and molecular evidence.  Bryologist 113: 149-163, f. 1-5.  J.J. Engel and X. He. 2010.

127.Austral Hepaticae 47. Plagiochila microdictyon Mitt. (Plagiochilaceae), a misunderstood species from New Zealand.  Bryologist 113: 34-38, f. 1-2.  J.J. Engel, and G. L. Merrill.  2010.

128.Early land plants today (ELLPT):  How many liverwort species are there?  Phytotaxa 9: 22-40.  von Konrat, M., L. Söderstrom, M. Renner, A. Hagborg, A. Briscoe and J. Engel.  2010.

129.Studies on Lophocoleaceae. XXII. The systematic position of Ammphilophocolea R.M.Schust together with comments on the status of Tetracymbaliella Grolle and Lamellocolea R.M.Schust.  Phytotaxa 9: 41-52, f. 1-4.  J.J. Engel, X. He and D. Glenny. 2010.

130.Early land plants today:  Taxonomy, systematics, and nomenclature of Gymnomitriaceae H. Klinggr.  Phytotaxa 11: 1-80.  Vána, J., L. Söderstrom, A. Hagborg, von Konrat, M., and J. Engel.  2010.

131.Studies on New Zealand Hepaticae.  39-55.  More new taxa, combinations, typifications and synonymy in Plagiochila from New Zealand (Plagiochilaceae).  Nova Hedwigia 91: 501-517.  J.J. Engel and G. L. Merrill.  2010.  (Nov.)

Professional affiliations

American Bryological and Lichenological Society

International Association of Bryologists